Discussion
Strength of social norms and associated norm-enforcement differ widely around the globe. The models and experiments presented in this paper demonstrate that high levels of threat create evolutionary pressures toward stronger norms for organizing social action, whereas low levels of threat allow for cultural populations with weaker norms. Cultural groups facing chronic high levels of threat evolve higher norm-adherence and higher levels of norm-enforcement than cultures exposed to relatively low levels of threat, both in cooperation as well as coordination games, and these results are mediated by the property of diminishing marginal returns in fitness of payoffs. Our models also show that temporary increases in threat cause norm strength to increase until the threat subsides, at which point behavioral norms begin returning to their previous level. This is analogous, for example, to situations where a society that has had relatively low levels of threat is faced with a temporary threat on its own soil (e.g., the 9/11 in the U.S.). Thus these results support correlational research that show that exposure to threat is an important driving factor in the evolution and maintenance of cultural differences in norm-strength and punishment of deviation (Gelfand et al., 2011; Harrington & Gelfand, 2014).